Not in a position to test atm, but the defs seem to imply some interaction between Animus stones and the tree similar to how braziers next to each other increase their ability. This needs to be investigated
Topic on Talk:Anima tree
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Just did, and indeed although the shrines don't have an effect the stones give +2% per stone up to 4 stones to shrines, anima tree or other stones.
I'll go add it in the article